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The correct understanding lies in the function of a template file within Revit. Template files are used to establish a baseline for projects by containing preset configurations, including families, views, and some settings. Consequently, the door and window sizes, which are part of the family definitions, are stored in the template file. When a new project is created using this template, these predefined sizes are readily accessible and can be utilized throughout the project, ensuring consistency and saving time.

While door and window families can be loaded or modified within a project, their initial settings are often set within the template for efficiency. This practice helps maintain standard dimensions and styles as the project evolves.

The other options do not provide the correct context regarding where this essential information is primarily stored. The project file directly incorporates various elements, including families that can be customized after the project starts. A separate configuration file is not typically part of the Revit organization and model settings refer more to graphical display than actual dimensional properties.